Holotype foot of Macrophalangia canadensis

Macrophalangia (meaning "large toes") is another theropod from the Red Deer Locality. First described by Charles Sternberg in 1932, Macrophalangia is known from a partial foot (see image), which looks very much like that of an ornithomimosaur. It has long, slender toes ending in flattened but quite pointed claws. However, unlike the three-toed ornithomimosaur foot, the foot of Macrophalangia has four toes: three long, slender ones and a smaller first toe typical of theropods. Does Macorphalangia represent a primitive type of ornithomimosaur? If so, then its appearance in late Cretaceous rocks is unexpected. Alternitavely, these remains may indicate a completely new group of slender-toed theropods in the late Cretaceous, the elmisauridae, containing the species Macrophalangia, Chirostenotes and Elmisaurus.

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